Sanger, who once described himself as "just a chap who messed about in his lab", worked with colleagues to develop a rapid method of DNA sequencing - a way to "read DNA" - which became the forerunner for the work on mapping the human genome.
He won his first Nobel Prize for Chemistry in 1958 for work on determining the structure of insulin, and the same Nobel 22 years later for his work on DNA, the material that carries all the information about how living things look and function.
Colin Blakemore, a Cambridge professor of neuro
"His invention of the two critical technical advances - for sequencing proteins and nucleic acids - opened up the fields of molecular biology, genetics and genomics," he said in a statement responding to news of Sanger's
(Reporting by Kate Kelland; editing by Patrick Graham)